$title$
$if(subtitle)$
$subtitle$
$endif$
$for(author)$
$if(author.last)$and $endif$
$author.name$$if(author.correspondence)$$if(author.email)$✉ $endif$$endif$$for(author.institute_indices)$$author.institute_indices$$sep$,$endfor$$if(author.equal_contributor)$,$if(equal_contributor_symbol)$$equal_contributor_symbol$$else$*$endif$$endif$$sep$,
$endfor$
$for(institute)$
$institute.index$$institute.name$$if(institute.address)$, $institute.address$$endif$
$endfor$
$if(has_equal_contributors)$
$if(equal_contributor_symbol)$$equal_contributor_symbol$$else$*$endif$These authors contributed equally to this work
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$if(has_correspondence)$
Correspondence: $for(author)$$if(author.correspondence)$$if(author.email)$$author.name$
<$author.email$> $endif$$endif$$endfor$
$endif$
$if(abstract)$$abstract$
$endif$
$if(doi)$doi: $doi$
$endif$
$for(include-before)$
$include-before$
$endfor$
$body$